The research team turned to MaxEnt, short for maximum entropy modelling, one of the most widely used tools in modern biogeography. The technique works by comparing the environmental conditions at locations where a species has been recorded with the conditions available across the whole study area, then estimating the probability of occurrence at every grid cell. Rather than requiring absence data, which is notoriously unreliable, MaxEnt builds a picture of a species’ ecological niche from presence records alone. For a country as topographically and climatically varied as Algeria, which spans Mediterranean coastline, high plateaus and the northern fringe of the Sahara, this approach is particularly powerful because it can capture the full breadth of conditions a species tolerates.

To drive the models, the researchers assembled occurrence records for both oak species, drawing on field knowledge and global biodiversity databases, and combined them with a suite of environmental predictor variables covering temperature, precipitation and terrain. Crucially, they projected the models forward using future climate scenarios for three time horizons, allowing them to track not just whether habitat will be lost or gained, but where the geography of suitability will shift. The performance of the models was assessed using the Area Under the Curve, or AUC, a standard statistical measure of discrimination. Values above roughly 0.8 are generally considered good, and above 0.9 excellent. Both oak models exceeded 0.96, a strikingly high score that indicates the models separated suitable from unsuitable terrain with near-perfect reliability.

The single most influential variable shaping the distribution of both species was the Martonne aridity index, a classic climatic measure that combines annual precipitation with mean temperature to express the degree of water availability across a landscape. Its dominance in the models makes ecological sense. Mediterranean oaks live at the dry edge of what trees can tolerate, and their growth, survival and regeneration are tightly governed by the balance between water arriving as rain and water lost to the atmosphere. In Algeria, where warming is outpacing the global average and droughts are becoming more frequent and severe, this finding underscores that aridity, not temperature alone, is the master variable controlling the fate of these forests.

When the researchers projected the models into the future, the two species told different stories. For cork oak, the annual rate of highly suitable habitat is expected to decline under every future scenario, with projected losses ranging from 0.040 percent to 0.191 percent per year. The numbers may look small, but compounded over decades they translate into a meaningful contraction of the species’ core range in Algeria. Cork oak is already under pressure across the western Mediterranean from land abandonment, overgrazing, fire and unexplained decline phenomena, so any additional climatic squeeze on its best habitat is cause for concern for foresters and conservationists alike.

Holm oak, by contrast, showed far greater stability. Its highly suitable habitat is projected to change only slightly, with shifts ranging from a loss of 0.021 percent to a gain of 0.164 percent per year depending on the scenario. This hardy evergreen is famously drought-tolerant, capable of surviving on thin soils and withstanding long summer dry seasons, and the model results reflect that toughness. In practical terms, the study suggests that holm oak could even hold its ground or modestly expand in some parts of Algeria as conditions change, making it a candidate species for stabilising forests in an increasingly arid landscape.

Perhaps the most vivid finding concerns elevation. The average elevation of suitable habitat is projected to shift under each climate scenario, ranging from 866 to 1,025 metres for cork oak and from 978 to 1,172.5 metres for holm oak. In other words, as the lowlands warm and dry, the climatic comfort zone for both species climbs the mountainsides, and holm oak consistently occupies higher ground than cork oak. This pattern echoes a well-documented global phenomenon in which mountain plant communities shift upslope in response to warming. The catch, as ecologists have repeatedly warned, is that mountains are cones: the higher a species climbs, the less land remains available, and slow-moving trees may lag far behind the pace of climate change.

The models also revealed a striking directional signature. Under most future scenarios, the highly suitable habitat for cork oak shifts towards the northeast of the country, while for holm oak it shifts towards the northwest. These compass bearings matter enormously for conservation planning. Northeastern Algeria, with the humid coastal ranges of Kabylia and El Kala, retains some of the wettest conditions in the country, and the projected northeastward drift of cork oak suitability suggests these areas will become increasingly critical refuges. The northwestward shift for holm oak points to a different set of priority landscapes, and the divergence between the two species means that a single, one-size-fits-all protected area network will not serve both trees equally well.

The practical value of this work lies in its maps. By identifying where habitat is highly suitable now and where it is likely to remain suitable through the end of the century, the study gives Algerian foresters a decision-support tool of real consequence. Rather than reacting to forest decline after it happens, managers can use these projections to prioritise which forest stands to protect, where to focus regeneration efforts, and which zones should be preserved as climate refugia. The authors emphasise that the maps are intended precisely for this purpose: helping foresters select the best conservation strategy for cork oak and holm oak and choose appropriate forest zones to be preserved before the climate window closes.

The study also sits within a growing body of Algerian research on climate-driven range shifts, including recent work on Atlas cedar, Aleppo pine, Phoenician juniper and the endemic oak Quercus afares, all pointing in a consistent direction: the country’s forest flora is being reshuffled by warming and drying. What makes the new findings compelling is the contrast between two closely related, sympatric oaks responding so differently to the same climatic pressures. As the Mediterranean basin warms faster than the global average, the fate of these forests will depend on whether conservation planning can anticipate the movement that the models now make visible. For Algeria’s cork and holm oaks, the future is not yet written, but the map of where it will unfold has finally been drawn.

Subject of Research: Species distribution modelling of cork oak and holm oak habitats in Algeria under climate change

Article Title: Modelling the current and future potential distributions of Quercus suber and Q. ilex (Fagaceae) in Algeria

Article References: Rais, H., Laala, A., Meghzili, I., Mennour, H., & Boucenna, H. (2026). Modelling the current and future potential distributions of Quercus suber and Q. ilex (Fagaceae) in Algeria. Plant Biosystems, 160(5), Article 239. https://doi.org/10.1007/s44473-026-00241-2

Image Credits: AI Generated

DOI: 10.1007/s44473-026-00241-2

Keywords: Quercus suber, Quercus ilex, Algeria, MaxEnt, species distribution modelling, climate change, aridity index, habitat suitability, Mediterranean forests, cork oak, holm oak, forest conservation

The team focused on three oak species that dominate the climax forests of acidic substrates across the peninsula: cork oak (Quercus suber), Pyrenean oak (Quercus pyrenaica) and pedunculate oak (Quercus robur). These species sit at different points along a moisture gradient. Pedunculate oak, a temperate and humidity-loving species, thrives in the cool Atlantic north. Pyrenean oak, a marcescent species that holds its withered leaves through winter, occupies a transitional submediterranean zone. Cork oak, the iconic evergreen of Mediterranean silviculture, tolerates pronounced summer drought. Because the three species often coexist in the same landscapes, distributed along gradients from rocky ridge tops to moist valley bottoms, the researchers reasoned that climate change could trigger a domino effect: as each species retreats from its drier margins, a more drought-adapted congener may move in to take its place.

To test this idea, the researchers built high-resolution species distribution models at a one-square-kilometre resolution, integrating nearly a quarter of a million cleaned occurrence records from GBIF, national biodiversity databases and expert-curated sources. For pedunculate oak, the team applied a spatial thinning procedure that reduced more than 205,000 records to about 27,000, mitigating oversampling while preserving the species’ environmental coverage. The models combined nineteen bioclimatic indices from the CHELSA dataset with soil pH, soil texture, a Topographic Ruggedness Index and a Topographic Wetness Index, all chosen through hypothesis-driven screening that excluded collinear predictors.

The modelling framework was an ensemble approach implemented in the biomod2 package in R, combining eight techniques ranging from generalized linear models and generalized additive models to random forests, artificial neural networks and maximum entropy. Each model was validated over ten rounds of cross-validation, and only algorithms exceeding a True Skill Statistic threshold of 0.7 were averaged into the final ensembles. Performance was strong across the board: TSS values exceeded 0.83 and AUC scores surpassed 0.97 for all three species, with cork oak models performing best. Future projections drew on five general circulation models selected for their strong performance over Europe, run under two Shared Socioeconomic Pathways: an intermediate SSP3-7.0 scenario and a pessimistic SSP5-8.5 scenario, for the mid-century period 2041 to 2070 and the late-century period 2071 to 2100.

The environmental variables that best explained each species’ current distribution revealed their distinct ecological thresholds. For pedunculate oak, the minimum temperature of the coldest month was the dominant predictor, reflecting its tolerance of subzero winters, followed by topographic ruggedness and precipitation of the warmest quarter. Pyrenean oak’s distribution hinged on annual precipitation and warm-season rainfall together with soil pH, underscoring its dependence on water availability during the hot, dry summer. Cork oak, by contrast, was governed primarily by soil pH and mean annual temperature, favouring sites where winter minima stay above freezing. All three species are strictly silicicole, confined to acidic substrates, which means that climatically suitable areas on limestone soils will remain off-limits no matter how the climate shifts.

The projected trajectories diverge sharply. Pedunculate oak’s current range is expected to remain largely stable, with between 73 and 84 percent of its present suitable area persisting across scenarios, though its total suitable area declines slightly as losses outweigh gains. Cork oak emerges as the clear winner: its suitable range expands in every scenario and time frame, gaining between 32 and 44 percent of new territory, pushing northward and inland, with potential altitudinal advances reaching as far as the Pyrenees, while losing almost nothing at its southern edge. Pyrenean oak fares worst. Its total suitable area shrinks in all scenarios, with losses of up to nearly 60 percent of its current range concentrated in the southern and inland parts of the peninsula, and only limited compensatory gains at higher elevations.

When the researchers overlaid the range dynamics of different species, the turnover patterns became strikingly clear. Wherever one species loses suitability while another gains or holds steady in the same grid cells, a potential replacement is flagged. The dominant signal is a turnover from Pyrenean oak to cork oak, projected across roughly 20 to 47 percent of Pyrenean oak’s currently suitable range depending on the scenario, intensifying toward the end of the century and under harsher emissions. This replacement is expected to begin at the species’ southern fringes and spread into the interior. A smaller turnover, from pedunculate oak to Pyrenean oak, is projected across about 5 to 8 percent of the former’s range, mostly in low-altitude inland areas at the southern edge of its distribution. In some coastal zones of western and northeastern Iberia, cork oak may ultimately replace both of the other species where they both become unsuitable.

A crucial dimension of the study is its analysis of protected areas. Between a quarter and nearly half of the projected turnover zones fall inside classified conservation sites, including Natura 2000 areas designated under the European Union’s Habitats Directive, national parks and nature reserves. The turnover from Pyrenean oak to cork oak carries a direct policy consequence: it implies a shift from habitat 9230, the Galicio-Portuguese oak woods, to habitat 9330, cork oak forests, within sites whose management plans were written for the habitats of the past. The researchers argue that conservation targets in these sites will need periodic revision to maintain habitat continuity under changing climates, and that protected areas can serve as demonstration cases for adaptive management on the privately owned lands that dominate the Portuguese landscape.

The authors are careful to frame these projections as shifts in potential climatic and edaphic suitability rather than deterministic predictions of stand-level replacement. The models do not explicitly simulate dispersal, disturbance regimes or biotic interactions, and oak species are notoriously slow to migrate relative to the pace of climate change. Yet the convergent trends across multiple climate models and scenarios lend weight to the central inference: that a gradual Mediterraneanization of the Iberian landscape is underway, marked by longer summer dry periods and reduced annual precipitation, in which drought-adapted oaks progressively occupy the space vacated by their moisture-dependent relatives.

The practical implications reach deep into forestry. The researchers recommend protecting natural regeneration of the species likely to dominate in the future, since cork oak individuals already occur near Pyrenean oak stands on rocky outcrops and within forests. Where natural regeneration is insufficient, they point to seed sowing and planting techniques, including autumn sowing to maximize root development before summer drought, selection of larger acorns, protection from seed predators such as wild boar, and the use of large containers to avoid root deformation. They also advocate multi-provenance strategies that draw reproductive material from southern or low-altitude populations already adapted to drier conditions, combined with mixed-species stands to hedge risk. If guided by such adaptive management, the authors conclude, species turnover need not spell the loss of Iberian forests; it may instead be the mechanism by which they retain canopy cover, ecological function and resilience through a century of rapid environmental change.

Subject of Research: Projected climate-driven species turnover among Iberian oak forests

Article Title: Exploring turnover dynamics in Iberian Oak forests under climate change scenarios

Article References: Passos, I., Figueiredo, A., Gonçalves, J., Ribeiro, M. M., & Vila-Viçosa, C. (2026). Exploring turnover dynamics in Iberian Oak forests under climate change scenarios. Discover Conservation, 3(1), Article 14. https://doi.org/10.1007/s44353-026-00084-0

Image Credits: AI Generated

DOI: 10.1007/s44353-026-00084-0

Keywords: climate change, oak forests, Quercus suber, Quercus pyrenaica, Quercus robur, species distribution models, species turnover, Iberian Peninsula, Mediterranean forests, forest management, conservation, habitat suitability